Breeding performance in the wild by Lear's Macaws

A researched paper entitled " Breeding to non-breeding population ratio and breeding performance of the globally endangered Lear's Macaw (Anodorhynchus leari); conservation and monitoring implications\" was published by Bird Conservation International in February 2014.  This highlighted the high performance of Lear's macaws in the wild in reproduction compared to other macaw species. I find this really interesting as Lear's macaws have proven to be quite prolific in breeding in captivity, e.g at Loro Parque in Tenerife. I was sent this paper recently by Acadaemia 

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Here is the paper by Erica.C.Pacifico, Eduardo. B.Barbosa, Thiago Filadelfo, Kleber. G.Oliveira, Luis.F.Silveira and Jose.L.Tella

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 " Naturam expellas furca, tamen usque recurret "

( If you drive out nature with a pitchfork, she will soon find a way back)

Horace (65-8 BC)